A shallow-water cyrtocrinid crinoid (Articulata) from the upper Albian of the Western Pyrenees, North Spain

The cyrtocrinid crinoid Proholopus holopiformis (Remeš, 1902) is described from the upper Albian succession that may be included in Albeniz unit to east of Iruñea-Pamplona (Navarre, Western Pyrenees, Spain). Although based on partially disarticulated material, this taxon preserve calyx, stem, attachment structure, and brachial plates arms. was previously Upper Jurassic-Lower Cretaceous Czech Republic Lower France Crimea, thus occurrence Spain represents youngest species. This allows a better characterization family Proholopodidae originally calyx morphology only expands its distribution Albian. inhabited fore-reef areas coral-sponge bioconstructions. Based coral types, crinoids are thought have dwelled near euphotic-oligophotic zones transition and, thus, thrive shallow depths. Most specimens bite marks compatible with cidaroid predation. one occurrences cyrtocrinids inhabiting marine environments before their migration deep sea due ongoing Mesozoic Marine Revolution. • A Spain. cups, stems, brachials, holdfast allowing well understanding morphology. Sedimentological analysis suggest P. lived water conditions less than 50 m deep. All analyzed show external predation by echinoids. environments.